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The California Academy of Sciences is a research institute and natural history museum in San Francisco, California, that is among the largest museums of natural history in the world, housing over 46 million specimens. The Academy began in 1853 as a learned society and still carries out a large amount of original research. It is California's oldest museum.

Completely rebuilt in 2008, the building covers 400,000 square feet (37,000 square metres).The primary building in Golden Gate Park reopened on September 27, 2008.

The California Academy of Sciences, California's oldest operating museum and research institution for the natural sciences, is governed by a forty-one member Board of Trustees who are nominated and chosen by the California Academy of Sciences Fellows. The Academy Fellows are, in turn, "[n]ominated by their colleagues and appointed by the Board of Trustees...the Fellows remain members of the Fellowship for life." The Board of Trustees are then responsible for appointing the executive management of the Academy, who in turn are responsible for overseeing the Academy's overall operation and the hiring of its other managers and employees.

Besides its function as source of public science education through its museum, the California Academy of Sciences also operates the Institute for Biodiversity Science and Sustainability (IBSS) as its research arm, contributing some of the world's most important research in the fields of taxonomy, phylogenetics, and biodiversity studies. Although one aspect of the IBSS is available for view by museum patrons at the science "project lab" exhibit, most of the research happens in laboratories and facilities "behind the scenes" and not observable by the public. In fact, unbeknownst to most patrons, research and administrative facilities occupy nearly 50% of the Academy's physical structure.


The main thrust of the exhibits is natural history. The main venues of the museum include the following:

Besides its museum programs, the California Academy of Sciences offers many educational and community outreach programs to members of the public at large.

Academy scientists, under the Academy's Institute for Biodiversity Science and Sustainability, conduct systematic and conservation research in several different fields, including anthropology, marine biology, botany, entomology, herpetology, ichthyology, invertebrate zoology, mammalogy, ornithology, geology, and paleontology. There also is a strong emphasis on environmental concerns, with all the various departments collaborating closely to focus on systematic biology and biodiversity. Academy researchers study life around the world: a 2011 expedition to the Philippines discovered an estimated 300 species new to science. The Academy publishes the peer-reviewed journal Proceedings of the California Academy of Sciences, as well as Occasional Papers, Memoirs, and Special Publications.

The California Academy of Natural Sciences was founded in 1853, only three years after California joined the United States, becoming the first society of its kind in the Western US. Its stated aim was to undertake "a thorough systematic survey of every portion of the State and the collection of a cabinet of her rare and rich productions." It was renamed as the more inclusive California Academy of Sciences in 1868.

The Academy had a forward-thinking direction to the involvement of women in science, passing a resolution in its first year of existence that the members "highly approve of the aid of females in every department of natural science, and invite their cooperation." This policy led to several women being hired into professional positions as botanists, entomologists, and other occupations during the 19th century, when opportunities for women in the sciences were limited, and often, those that existed were restricted to menial cataloging and calculation work. In 1892, Alice Eastwood, a botanist, was hired by the Academy and worked there until she retired in 1949. She created the collection of rare plants, which was saved when the Academy was destroyed in the 1906 San Francisco earthquake

The Academy's first official museum opened in 1874 at the corner of California and Dupont Streets (now Grant Avenue) in what is now Chinatown, and drew up to 80,000 visitors a year. To accommodate its increasing popularity, the Academy moved to a new and larger building on Market Street in 1891, funded by the legacy of James Lick, a 19th-century San Francisco real estate mogul, entrepreneur, and philanthropist.

However, only fifteen years later, the Market Street facility fell victim to the 1906 San Francisco earthquake and three days of fire, which also wiped out all but a wheelbarrow full of the Academy's library and specimen collections. In the widespread destruction occurring in the aftermath of the quake, Academy curators and staffers only were able to retrieve a single cart of materials, including Academy minute books, membership records, and 2,000 type specimens. The 1905-1906 scientific collecting expedition to the Galápagos Islands (the first of several sponsored by the academy to the archipelago) already was underway, and it returned seven months later, providing replacement collections for those lost.

In 1916, the Academy moved to the North American Hall of Birds and Mammals in Golden Gate Park, the first building on the site that was to become its permanent home. In 1923, the Steinhart Aquarium was added, followed in 1934 by the Simson African Hall.

During World War II, the Academy contributed to the American war effort by using its workshop facilities to repair optical and navigational equipment for United States Navy ships; San Francisco was a major port for the Pacific War arena.

The post-war years saw a flurry of new construction on the site; the Science Hall was added in 1951, followed by the Morrison Planetarium in 1952. The Morrison Planetarium was the seventh major planetarium to open in the United States and featured a one-of-a-kind star projector, built by Academy staff members (in part using the expertise gained doing the optical work for the US Navy during World War II). The Academy Projector projected irregularly shaped stars, rather than the circular stars projected by many optical star projectors. The irregular shapes were created by placing variously sized grains of silicon carbide onto the glass star plates by hand, then aluminizing the plates, and brushing away the silicon carbide grains.

In 1959, the Malliard Library, Eastwood Hall of Botany, and Livermore Room all were added. Throughout the 1960s, universities concentrating on the new field of molecular biology divested themselves of their traditional specimen collections, entrusting them to the Academy and leading to a rapid growth of the Academy's holdings.

In 1969, another new building, Cowell Hall, was added to the site. In 1976, several new galleries were opened, and the following year, in 1977, the "fish roundabout" was constructed.

Prior to the old building being torn down in 2005, there was a Life through Time gallery, housing a large display on evolution and paleontology. There was a Gem and Mineral Hall, a section on Earthquakes, and a Gary Larson exhibit.

The Academy buildings were damaged significantly in the 1989 Loma Prieta earthquake. Subsequently, the Bird Hall building was closed to ensure public safety. The inadequately engineered Steinhart Aquarium suffered dramatic seismic damage from the 1989 earthquake, as well.

As plans were made to repair the damage and make the buildings seismically stable, it was realized that a considerable amount of work would be needed to bring the buildings up to modern standards. This led to the idea of giving the Academy a complete overhaul, thus motivating the closing of the main site.

Construction began on the new $500 million building on September 12, 2005, while the exhibits were moved to 875 Howard Street for a temporary museum.

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